5. Jenkins gave the writer a selfed stock homozygous for mottled aleurone. He had found it extremely difficult to get a homozygous stock in which all seeds showed mottling since the mottled condition is extremely susceptible to the action of modifiers. This strain was turned over to the writer because it seemed possible that this case was similar to the a-Dt situation where one gene stimulates the mutability of another. The mottling proved to be caused by an r allele and was not due to another gene causing r to mutate. This allele is a new member of the R series. The mottled condition resembles most closely that produced by a single dose of R. It is not the same as the marbled and stippled alleles found in certain strains from Mexico and South America.

M. M. Rhoades