1. In Garrapata corn from the Province of Salta in Argentina and from Bolivia, spotted aleurone is due to a dominant r modifier giving mottled aleurone.

Mottled x a and c testers gives self color

Mottled x r testers gives mottled F1, but in F2 some colorless kernels appear. The modifier is independent from pr and from a and c but seems to be linked with r. The r modifier is designated Mr. The backcross: r Mr Pr/r mr pr x r mr pr gave
Mottled purple66
Mottled red59

[Mr has been used by Kvakan for midrib (Linkage Summary, p. 15) but the stock has been lost. Seeds sent look like "stippled", which is either an allelomorph of r or very closely linked with it.]